What resolution did you have the iMac setup when you were having issues? Even though you are emulating 2560x1440, LR is retina aware, and utilizes the full resolution of the display in the image areas of the program, so if the iMac is choking on the massive amount of pixels in the image area, changing to an emulated resolution shouldn’t make much of a difference. Emulating a lower working resolution is one of the basic concepts behind the design, it removes the need for using a “native” resolution.
I have the resolution set to "Best for retina", which is the 2560x1440 high res mode. At this resolution, Lightroom has to work with a very large number of pixels. I have three tests that I think demonstrate that it is in fact this large number of pixels that is affecting performance:
1. If I zoom in to 2:1, adjustments smooth out.
2. If I resize the the Lightroom window to fill about 1/3 of the screen, adjustments likewise get much smoother.
3. If I put the screen in 2560x1440 low resolution mode, which turns it essentially into a standard 27-inch iMac display, again adjustments are smooth.
In all of those cases Lightroom is working with fewer pixels and performance improves. Having said all of that, I'm now OK with performance. Whether it's something I did when I was trying various things to work through the issue, or perhaps just a matter of me adjusting to how it behaves, in practice the only place that I see really obvious jerkiness is with the graduated filter and transformations. I rarely use those. And there's some funkiness with cropping, but I think that can be fixed by Adobe.