Why those two cameras? I can see one camera for the landscape opportunities (either the 645z OR the Sony A7R II), and another camera for the long lens action. If I were going, I would be taking a Canon 7D2 with its high burst rate, generous RAW buffer, sturdy build, and crop factor convenience giving the same field of view for 400mm on crop as you have for 600mm on full frame. I would mount a Canon 100-400mm f/variable L IS II (brand new) for hand-held shots or Canon 200-400mm f/4 L IS with built-in 1.4x teleconverter for tripod/monopod/vehicle - mounted shooting. If money is no object and you are experienced at bird or wildlife or sports photography and plan to use tripod/monopod/vehicle mount, and you want best low light action performance, a Canon 1DX full frame camera plus the 200-400 would be ideal. If you are a newbie to wildlife, the 7D2 and 100-400 II would be a lightweight and easier to handle combo. ++++++ Practice+++++++ on local birds, dogs, racecars, whatever to familiarize yourself with the complex autofocus options of either of the Canon cameras and get warmed up on your panning skills. Don't go on safari expecting to pick up skills. I wouldn't expect to see the kind of high speed, high accuracy supertelephoto performance from Sony A7R II plus Metabones adapter plus Canon lens that you would get from the Canon camera plus Canon lens, no matter what Sony claims. This is a demanding AF use and not the sort of situation that Sony has been using for demonstration. The supertelephoto motors are driven from the camera's battery, speed depends somewhat on the voltage of the battery. Also - you could rent the supertelephoto.
My situation:stills shooter, small woman, currently shoots wildlife and birds (in flight) with 2 kg combo handheld Canon 400mm f/5.6 and APS-C camera Canon 60D, shortly to upgrade to 7D2.